Can You Use a PCIe Express Card In a Slot?

What is the difference between PCIe Gen 3 and PCIe Gen 4? m2 pcie 4 0 in 3 0 slot
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m2 sata slot My questions are: 1 Is the PCI-E NVME SSD compatible with my Z390 Godlike MOBO which has PCI-E slots? 2